The largest mIMR in the nucleocapsid spans the two Cys-His boxes [NCBI:1F6U ]. Figure 3A illustrates the largest mIMR in the nucleocapsid – #6-gag. This mIMR spans both zinc knuckles and the spacer between them. Each of the next largest mIMRs in the NC, translates one of the Cys-His boxes. Figure 3B illustrates the first Cys-His box. Figure C (same polar orientation as A and B, but rotated) illustrates the two longest rdIMRs in Gag that occur in the nucleocapsid – $1-gag and $4-gag – which overlap; within the overlap region (in purple) two amino acids bind the zinc ion .